Let me try and put this in a different perspective.
The plane relies on thrust, through the form of displacing air, to move. Displacing air, has nothing to do with the ground. Right.
Its wheels are there to minimize friction between it and the ground.
Now, replace the airplane with a skatebaord, kinda like Billabond did. It's much like the airplane, only it lacks something to make it move.
And now, replace the runway with a treadmill...pointing almost straight down, with the conveyor moving upwards. That way, the skateboard has something to propel it, namely, gravity, which is also independent of air. If you set the skateboard on it and program it to increase its speed to match teh skateboard's while the skateboard accellerates, it won't stop it.
The wheels can not generate enough friction to counteract the force causing it to accellerate...the plane is designed so that the wheels do not have enough friction to stop it. If you had the treadmill spinning at 30mph, several times the skateboard's downward velocity, the skateboard would still fall.
Thus, the skateboard is moving relative to the air, as would the plane, because there is not enough friction in the wheels to keep it from moving.